A United States fighter jet shot down an armed pro-regime drone in southern Syria on Tuesday, according to us -led coalition officials.
USA anti-ISIS strategy in Syria relies on the Syrian Democratic Forces (SDF), a group largely composed of Kurdish fighters.
A U.S. F-15E Strike Eagle shot down the Shaheed-129 UAV at approximately 12:30 a.m.
The official added the US has "repositioned" its air forces in Syria in the wake of the USA downing of the Russian-made Syrian jet.
Moscow expects "detailed" explanations from Washington after the USA on Sunday shot down a Syrian drone bomber, Russian Foreign Minister Sergey Lavrov said on Tuesday.
Coalition forces are using the Al-Tanaf base by Syria s borders with Jordan and Iraq to train anti-jihadist Syrian fighters and stage attacks against IS.
On the other hand, Russia, along with Iran, has been assisting the Syrian government in its fight against terrorist groups wreaking havoc in the Arab country since 2011. Its defense ministry said it plans to track joint coalition aircraft west of the Euphrates River and treat them as targets, officials said Monday.
Moscow said it was suspending an incident hotline set up two years ago and warned that it would consider worldwide aircraft operating in central Syria "aerial targets".
It was shot down near Tanf, an outpost of the US-led coalition.
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"We are very well prepared and positioned to defend ourselves", Operation Inherent Resolve spokesman Army Col. Ryan Dillon said earlier this month. And on Monday, a US fighter jet shot down a pro-Syrian regime drone in southeastern Syria.
Tuesday's statement from the worldwide coalition stressed that the global effort remained focused on ISIS but said that coalition forces had the right to self-defense.
The U.S. could increasingly find itself engaging Syrian regime, Iranian, or even Russian military assets as the campaign to retake the Islamic State's capital of Raqqa escalates.
"This is not the time for the coalition to be in disarray", Dr Lina Khatib, head of the Middle East and North Africa Programme at Chatham House, an worldwide affairs think tank, told CNN. One of the officials said that UAV "was assessed as a threat".
"The coalition presence in Syria addresses the imminent threat ISIS in Syria poses globally".
"The US position militarily is weaker and any sign of a lack of unity within the worldwide anti-ISIS coalition can only be good news for Russian Federation and the Syrian regime". The US has defended its attack on the Syrian jet, stating that it was in defence of US forces seeking to capture Raqqa from Daesh (Islamic State). The force will continue normal operations in Iraq, reports Australia's News.com.
The shooting down underscores the growing tensions in the region as a battle develops for the control of eastern Syria, BBC Defence and Diplomatic Correspondent Jonathan Marcus says.
On Monday, the Iranian Revolutionary Guard Corps demonstrated some of its military prowess by launching ballistic missile strikes from western Iran against ISIS targets in the Syrian city of Deir ez-Zour, according to The Guardian.
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